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From Flat Foot to Fat Foot: Structure, Ontogeny, Function, and Evolution of Elephant “Sixth Toes”

via Nature: “Even though an elephant’s leg looks like a solid column, it actually stands on tip-toe like a horse or a dog. Its heel rests on a large pad of fat that gives it a flat-footed appearance. The pad hides a sixth toe — a backward-pointing strut that evolved from one of their sesamoids, a set of small tendon-anchoring bones in the animal’s ankle. This extra digit, between 5 and 10 centimetres long, had been dismissed as an irrelevant piece of cartilage. Almost 300 years after it was first described, Hutchinson finally confirmed that it is a true bone that supports the squishy back of the elephant’s foot. The ones on the hindfeet even seem to have joints.” The full-text is available to subscribers, Hutchinson, J. R. et al. Science 334, 1699–1703(2011).”

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